Lithology
The lower section of the formation is around 50–130 m thick, and consists of light grey sandstones with gravel, siltstone and rare coal beds.[3] At the Dubinino locality, the 50 m exposed section of the upper part of the formation shows an irregular rhythmic alteration of fine grained sandstone, siltstone and mudstone and coal seams typically a few tens of cm's but up to several meters thick.[1] The deposit is located on the South Eastern margin of the West Siberian basin[4]
Locality
Most of the fossils were found in the overburden of Berezovsk coal mine[ru], which is located in southern Krasnoyarsk Krai (Sharypovsky District) near the border with Kemerovo Oblast. The deposit is stratigraphically located in the upper member of the formation. The fossiliferous level of the locality is located above thick (> 50 m) coal seams and consists of unconsolidated silt and sand, which were deposited on an alluvial plain. Due to the fluvial origin of the sediment the remains are disarticulated and often are water worn, though they are mostly well preserved, which suggests they had not been significantly transported.[5] The fossils were largely obtained by screenwashing of the debris. 10-15 tons of material have been processed so far.
Paleobiota
Taken from[2] unless otherwise noted.
Fish
Fish of the Itat Formation
|
Genus
|
Species
|
Location
|
Stratigraphic position
|
Abundance
|
Notes
|
Images
|
Hybodus
|
Indeterminate
|
Berezovsk coal mine
|
Upper Member
|
Partial tooth
|
|
|
Acipenseriformes
|
Indeterminate
|
Berezovsk coal mine
|
Upper Member
|
Scales
|
|
|
Amiiformes
|
Indeterminate
|
Berezovsk coal mine
|
Upper Member
|
"Abundant vertebrae, jaws and vomer fragments, as well as isolated teeth, and numerous isolated rhomboidal scales"
|
|
|
Dipnoi
|
Indeterminate
|
Berezovsk coal mine
|
Upper Member
|
One complete upper tooth plate, several tooth plate fragments
|
|
|
Palaeonisciformes
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Undescribed
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Berezovsk coal mine
|
Upper Member
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Many partial skeletons
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Similar to Palaeoniscinotus and Pteroniscus
|
|
Amphibians
Amphibians of the Itat Formation
|
Genus
|
Species
|
Location
|
Stratigraphic position
|
Abundance
|
Notes
|
Images
|
?Eodiscoglossus[6]
|
Indeterminate
|
Berezovsk coal mine
|
Upper Member
|
Atlantal centrum
|
|
Frog
|
Kiyatriton[7]
|
K. krasnolutskii
|
Berezovsk coal mine
|
Upper Member
|
Dentary, atlantes, trunk vertebrae and fragmentary cranial and postcranial bones
|
Formerly known as "Berezovsk salamander B"; genus also known from Early Cretaceous Ilek Formation.
|
Crown group Salamander
|
Urupia[8]
|
U. monstrosa
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Berezovsk coal mine
|
Upper Member
|
Dentary, femur, an incomplete atlas, an anterior fragment of the left dentary, and fragments of trunk vertebrae
|
Estimated length of 480-560 centimetres
|
Stem-Salamander
|
Egoria [9]
|
E. malashichevi
|
Berezovsk coal mine
|
Upper Member
|
Atlantal centra, trunk vertebrae
|
Formerly known as "Berezovsk salamander A" Possibly neotenic
|
Stem-Salamander
|
Turtles
Turtles of the Itat Formation
|
Genus
|
Species
|
Location
|
Stratigraphic position
|
Abundance
|
Notes
|
Images
|
Annemys[10]
|
Indeterminate
|
Berezovsk coal mine
|
Upper Member
|
Thousands of isolated bones and several fragmented specimens, including three partial braincases and larger fragments of connected shell parts
|
Xinjiangchelyid
|
|
Testudines
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Indeterminate
|
Berezovsk coal mine
|
Upper Member
|
|
3 distinct taxa
|
|
Lepidosauromorphs
Lepidosauromorphs of the Itat Formation
|
Genus
|
Species
|
Location
|
Stratigraphic position
|
Abundance
|
Notes
|
Images
|
Lepidosauromorpha
|
Indeterminate
|
Berezovsk coal mine
|
Upper Member
|
Several fragments of dentaries and maxillae
|
Basal form similar to Marmoretta
|
|
Scincomorpha
|
Indeterminate
|
Berezovsk coal mine
|
Upper Member
|
Fused premaxillae and two dentary fragments additionally several fragmentary maxillae and dentaries
|
Two distinct taxa, one of which is two times smaller than the other.
|
|
Choristoderes
Choristoderes of the Itat Formation
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Genus
|
Species
|
Location
|
Stratigraphic position
|
Abundance
|
Notes
|
Images
|
Cteniogenys
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Indeterminate
|
Berezovsk coal mine
|
Upper Member
|
Skull roof bones, maxillary and dentary fragments, vertebrae, ribs
|
|
|
Pterosaurs
Pterosaurs of the Itat Formation
|
Genus
|
Species
|
Location
|
Stratigraphic position
|
Abundance
|
Notes
|
Images
|
?Rhamphorhynchidae
|
Indeterminate
|
Berezovsk coal mine
|
Upper Member
|
Numerous isolated teeth and very rare bone fragments
|
|
|
Crocodyliformes of the Itat Formation
|
Genus
|
Species
|
Location
|
Stratigraphic position
|
Abundance
|
Notes
|
Images
|
Goniopholididae[11]
|
Indeterminate
|
Berezovsk coal mine
|
Upper Member
|
Several dozens of isolated teeth, osteoderms, and a few cranial fragments
|
|
|
Non-avian dinosaurs
Non Avian DInosaurs of the Itat Formation
|
Genus
|
Species
|
Location
|
Stratigraphic position
|
Abundance
|
Notes
|
Images
|
Kileskus[12]
|
K. aristotocus
|
Berezovsk coal mine
|
Upper Member
|
Holotype description: "premaxilla, maxilla, a mandible fragment, and several limb bone parts"; referred material includes cervical and caudal vertebrae and numerous isolated teeth, as well as a fibula[13]
|
Proceratosaurid
|
|
Heterodontosauridae
|
Indeterminate
|
Berezovsk coal mine
|
Upper Member
|
Isolated maxillary and dentary teeth
|
|
|
Mamenchisauridae[14]
|
Indeterminate
|
Berezovsk coal mine
|
Upper Member
|
Numerous isolated teeth and two platycoelous posterior caudal vertebrae
|
Sauropod
|
|
Stegosauria[15]
|
Indeterminate
|
Berezovsk coal mine
|
Upper Member
|
Fragmentary skeletons of at least two individuals
|
|
|
Tritylodontids
Tritylodontids of the Itat Formation
|
Genus
|
Species
|
Location
|
Stratigraphic position
|
Abundance
|
Notes
|
Images
|
Stereognathus[16]
|
Indeterminate
|
Berezovsk coal mine
|
Upper Member
|
Upper and lower postcanine teeth
|
|
|
Insects
Numerous insect species are known from the Kubekovo village locality located within the upper member of the formation.[25]
Flora
See also
References
- ^ a b c Ivantsov, S. V.; Bystritskaya, L. I.; Krasnolutskii, S. A.; Lyalyuk, K. P.; Frolov, A. O.; Alekseev, A. S. (September 2016). "Middle Jurassic continental biota and paleolandscape in the Dubinino locality (Sharypovo area, Krasnoyarsk krai)". Stratigraphy and Geological Correlation. 24 (5): 486–502. doi:10.1134/s0869593816050038. ISSN 0869-5938.
- ^ a b "Middle Jurassic vertebrate assemblage of Berezovsk coal mine in western Siberia (Russia)". Global Geology. 19 (4): 187–204. 2016. doi:10.3969/j.issn.1673-9736.2016.04.01.
- ^ "THE ILANSK FORMATION AND ITS STRATOȚYPE". Russian Geology and Geophysics.
- ^ Le Heron, Daniel Paul; Buslov, Micha M.; Davies, Clare; Richards, Keith; Safonova, Inna (July 2008). "Evolution of Mesozoic fluvial systems along the SE flank of the West Siberian Basin, Russia". Sedimentary Geology. 208 (1–2): 45–60. doi:10.1016/j.sedgeo.2008.05.001.
- ^ a b c Averianov, Alexander O.; Martin, Thomas; Lopatin, Alexey V.; Schultz, Julia A.; Schellhorn, Rico; Krasnolutskii, Sergei; Skutschas, Pavel; Ivantsov, Stepan (2020-04-30). Hautier, Lionel (ed.). "Multituberculate mammals from the Middle Jurassic of Western Siberia, Russia, and the origin of Multituberculata". Papers in Palaeontology: spp2.1317. doi:10.1002/spp2.1317. ISSN 2056-2802.
- ^ Skutschas, Pavel P.; Martin, Thomas; Krasnolutskii, Sergei A. (2016-02-24). "First Jurassic frog from Siberia, Russia". Journal of Vertebrate Paleontology. 36 (3): e1101376. doi:10.1080/02724634.2016.1101376. ISSN 0272-4634.
- ^ Skutschas, Pavel P. (2015-10-28). "A new crown-group salamander from the Middle Jurassic of Western Siberia, Russia". Palaeobiodiversity and Palaeoenvironments. 96 (1): 41–48. doi:10.1007/s12549-015-0216-x. ISSN 1867-1594.
- ^ P. P. Skutschas & S. A. Krasnolutskii (2011). "A new genus and species of basal salamanders from the Middle Jurassic of western Siberia, Russia" (PDF). Proceedings of the Zoological Institute RAS. 315 (2): 167–175.
- ^ Skutschas, Pavel; Kolchanov, Veniamin; Krasnolutskii, Sergey; Averianov, Alexander; Schellhorn, Rico; Schultz, Julia; Martin, Thomas (2020-02-19). Porro, Laura Beatriz (ed.). "A new small-sized stem salamander from the Middle Jurassic of Western Siberia, Russia". PLOS ONE. 15 (2): e0228610. doi:10.1371/journal.pone.0228610. ISSN 1932-6203.
{{cite journal}}
: CS1 maint: unflagged free DOI (link)
- ^ Danilov, I. G.; Obraztsova, E. M.; Boitsova, E. A.; Skutschas, P. P. (March 2018). "Diversity of Middle Jurassic Turtles from the Berezovsk Quarry Locality, Krasnoyarsk Region, Russia: Morphology and Histological Study". Paleontological Journal. 52 (2): 188–200. doi:10.1134/s0031030118010070. ISSN 0031-0301.
- ^ Kuzmin, I.T.; Skutschas, P.P.; Grigorieva, O.I.; Krasnolutskii, S.A. (2013). "Goniopholidid crocodylomorph from the Middle Jurassic Berezovsk Quarry locality ( Western Siberia, Russia)". Proceedings of the Zoological Institute of the Russian Academy of Sciences. 317 (4): 452–458.
- ^ Averianov, A. O.; Krasnolutskii, S. A.; Ivantsov, S. V. (2010). "A new basal coelurosaur (Dinosauria: Theropoda) from the Middle Jurassic of Siberia". Proceedings of the Zoological Institute. 314 (1): 42–57.
- ^ O. Averianov, Alexander; Osochnikova, Anastasia; Skutschas, Pavel; Krasnolutskii, Sergei; Schellhorn, Rico; A. Schultz, Julia; Martin, Thomas (2019-09-16). "New data on the tyrannosauroid dinosaur Kileskus from the Middle Jurassic of Siberia, Russia". Historical Biology: 1–7. doi:10.1080/08912963.2019.1666839. ISSN 0891-2963.
- ^ Averianov, Alexander; Krasnolutskii, Sergei; Ivantsov, Stepan; Skutschas, Pavel; Schellhorn, Rico; Schultz, Julia; Martin, Thomas (2019-03-01). "Sauropod remains from the Middle Jurassic Itat Formation of West Siberia, Russia". PalZ. 93 (4): 691–701. doi:10.1007/s12542-018-00445-8. ISSN 0031-0220.
- ^ Averianov, A.O.; Krasnolutskii, S.A. (2009). "Stegosaur remains from the Middle Jurassic of West Siberia". Proceedings of the Zoological Institute RAS. 313 (2): 153–167.
- ^ Averianov, Alexander O.; Martin, Thomas; Lopatin, Alexey V.; Schultz, Julia A.; Skutschas, Pavel P.; Rico, Schellhorn; Krasnolutskii, Sergei A. (2017-09-03). "A tritylodontid synapsid from the Middle Jurassic of Siberia and the taxonomy of derived tritylodontids". Journal of Vertebrate Paleontology. 37 (5): e1363767. doi:10.1080/02724634.2017.1363767. ISSN 0272-4634.
- ^ Lopatin, A. V.; Averianov, A. O. (December 2007). "The earliest Asiatic pretribosphenic mammal (Cladotheria, Amphitheriidae) from the Middle Jurassic of Siberia". Doklady Biological Sciences. 417 (1): 432–434. doi:10.1134/s0012496607060063. ISSN 0012-4966.
- ^ Averianov, Alexander; Martin, Thomas; Lopatin, Alexey; Krasnolutskii, Sergei (2013-12-22). "Stem therian mammal Amphibetulimus from the Middle Jurassic of Siberia". Paläontologische Zeitschrift. 89 (2): 197–206. doi:10.1007/s12542-013-0217-x. ISSN 0031-0220.
- ^ Averianov, Alexander O.; Martin, Thomas; Lopatin, Alexey (2014-06-07). "The oldest dryolestid mammal from the Middle Jurassic of Siberia". Journal of Vertebrate Paleontology. 34 (4): 924–931. doi:10.1080/02724634.2014.837471. ISSN 0272-4634.
- ^ a b "New docodontians from the Middle Jurassic of Siberia and reanalysis of docodonta interrelationships" (PDF). Proceedings of the Zoological Institute RAS. 314 (2): 121–148. 2010.
- ^ Lopatin, A. V.; Averianov, A. O. (November 2005). "A New Docodont (Docodonta, Mammalia) from the Middle Jurassic of Siberia". Doklady Biological Sciences. 405 (1–6): 434–436. doi:10.1007/s10630-005-0158-y. ISSN 0012-4966. PMID 16485637.
- ^ a b c Averianov, Alexander O.; Martin, Thomas; Lopatin, Alexey V.; Schultz, Julia A.; Schellhorn, Rico; Krasnolutskii, Sergei; Skutschas, Pavel; Ivantsov, Stepan (2019-11-05). "Haramiyidan mammals from the Middle Jurassic of Western Siberia, Russia. Part 1: Shenshouidae and Maiopatagium". Journal of Vertebrate Paleontology. 39: e1669159. doi:10.1080/02724634.2019.1669159. ISSN 0272-4634.
- ^ A. O. Averianov, A. V. Lopatin and S. A. Krasnolutskii (2011). "The first Haramiyid (Mammalia, Allotheria) from the Jurassic of Russia". Doklady Biological Sciences. 437 (1): 103–106. doi:10.1134/S0012496611020074. PMID 21562957.
- ^ Averianov, A. O.; Lopatin, A. V.; Krasnolutskii, S. A. (2008). "An amphilestid grade eutriconodontan from the Middle Jurassic of Russia". Russian Journal of Theriology. 7 (1): 1–4. doi:10.15298/rusjtheriol.07.1.01.
- ^ "Kubekovo village (PIN collection 1255)". Paleobiology Database.